| Source of variation | Sum of squares | df | Mean square | F | P |
| Intersection | 4741.046 | 1 | 4741.046 | 533.987 | 0.000 |
| Site | 206.907 | 1 | 206.907 | 23.304 | 0.000 |
| Species | 875.159 | 4 | 218.790 | 24.642 | 0.000 |
| Site * Species | 303.467 | 4 | 75.867 | 8.545 | 0.000 |